Sabado, Oktubre 15, 2011
Science vs the church
Modern caricature of Galileo vs the church
Galileo vs the church/pope
Poor Galileo Galilei has become the poster child for the whole "small-minded church persecutes obviously correct scientist who threatens their power" thing. [Science Channel]
Galileo is in prison in his own house just because of assuming that the sun and not the earth is the center of the universe. According to the bible the earth is created first before the sun, so it appears to be the earth and not the sun is the center of the universe.
When Galileo invented the first telescope he discover a strong proof in the heliocentrism (sun center), because of the strong proof in heliocentrism, the pope in that time accuse Galileo to house arrest.
Copernicus vs the church
Before Galileo.Nicolaus Copernicus begun to make astronomy as a hobby, and he built a mini- observatory in their monastery. In that monastery he again prove the heliocentrism using trigonometry. But the consequences is too fast, the church began to consider him HERETIC.
Darwin against the creationism
According to the bible, God created us the way God look-like, but it has no proof.
If the bible has no proof, Charles Darwin has! Charles Darwin is a naturalist that study the life in the Galapagos island including finches. He use some specimens he collect from the island to prove and think about evolution. His Natural Selection was published in his book On the Origin of Species by Means of Natural Selection by the help of Alfred R. Wallace.
The church should be happy that our brightest figures-- the scientists are appreciating God's creation.
Biyernes, Oktubre 14, 2011
Sequential hermaphroditism
Sequential hermaphroditism (called dichogamy in botany) is a type of hermaphroditism that occurs in many fish, gastropods and plants. Here, the individual is born one sex and changes sex at some point in their life. They can change from a male to female (protandry), or from female to male (protogyny).[1] Despite which sex the organism changes to, those that change gonodal sex can have both female and male germ cells in the gonads or can change from one complete gonodal type to the other during their last life stage.[2]
Other examples of protandrous animals include:
In the California Sheephead (Pimelometopon pulchrum), a type of wrasse, when the female changes to male, the ovaries degenerate and spermatogenic crypts appear in the gonads. The general structure of the gonads remains ovarian after the transformation and the sperm is transported through a series of ducts on the periphery of the gonad and oviduct. Here sex change is age dependant. For example, the California sheephead stays a female for four years before changing sex.[7]
Other examples of protogynous organisms include:
In most ectotherms body size and female fecundity are positively correlated.[1] This supports Ghiselin’s size-advantage model, which is still widely accepted today. Kazancioglu and Alonzo (2010) performed the first comparative analysis of sex change in Labridae. Their analysis supports the size-advantage model by Ghiselin and suggest that sequential hermaphroditism is correlated to the size-advantage. They determined that dioecy was less likely to occur when the size advantage is stronger than other advantages[10]
Warner suggests that selection for protandry may occur in populations where female fecundity is augmented with age and individuals mate randomly. Selection for protogyny may occur where there are traits in the population that depress male fecundity at early ages (territoriality, mate selection or inexperience) and when female fecundity is decreased with age, the latter seems to be rare in the field.[1] An example of territoriality favoring protogyny occurs when there is a need to protect their habitat and being a large male is advantageous for this purpose. In the mating aspect, a large male has a higher chance of mating, while this has no effect on the female mating fitness.[10] Thus, he suggests that female fecundity has more impact on sequential hermaphroditism that the age structures of the population.[1]
The size-advantage model predicts that sex change would only be absent if the relationship between size/age with reproductive potential is identical in both sexes. With this prediction one would assume that hermaphroditism is very common, but this is not the case. Sequential hermaphroditism is very rare and according to scientists this is due to some cost that decreases fitness in sex changers as opposed to those who don’t change sex. Kazanciglu and Alonzo confirmed this in 2009. They found that the costs of changing sex only favored dioecy when the cost was very large but that some groups favored hermaphroditism. This indicates that the cost of sex change does not explain the rarity of sequential hermaphroditism by itself.[11]
Historically, dichogamy has been regarded as a mechanism for reducing inbreeding (e.g., Darwin, 1862). However, a survey of the angiosperms found that self-incompatible (SI) plants, which are incapable of inbreeding, were as likely to be dichogamous as were self-compatible (SC) plants (Bertin, 1993). This finding led to a reinterpretation of dichogamy as a more general mechanism for reducing the impact of pollen-pistil interference on pollen import and export (reviewed in Lloyd & Webb, 1986; Barrett, 2002). Unlike the inbreeding-avoidance hypothesis, which focused on female function, this interference-avoidance hypothesis considers both gender functions.
In many hermaphroditic species, the close physical proximity of anthers and stigma makes interference unavoidable, either within a flower or between flowers on an inflorescence. Within-flower interference, which occurs when either the pistil interrupts pollen removal or the anthers prevent pollen deposition, can result in autonomous or facilitated self-pollination (Lloyd & Webb, 1986; Lloyd & Schoen, 1992). Between-flower interference results from similar mechanisms, except that the interfering structures occur on different flowers within the same inflorescence and it requires pollinator activity. This results in geitonogamous pollination, the transfer of pollen between flowers of the same individual (Lloyd & Schoen, 1992; de Jong et al., 1993). In contrast to within-flower interference, geitonogamy necessarily involves the same processes as outcrossing: pollinator attraction, reward provisioning, and pollen removal. Therefore, between-flower interference not only carries the cost of self-fertilization (inbreeding depression; Charlesworth & Charlesworth, 1987; Husband & Schemske, 1996), but also reduces the amount of pollen available for export (so-called "pollen discounting"; Harder & Wilson, 1998]). Because pollen discounting diminishes outcross siring success, interference avoidance may be an important evolutionary force in floral biology (Harder & Barrett, 1995, 1996; Harder & Wilson, 1998; Barrett, 2002).
Dichogamy may reduce between-flower interference by minimizing the temporal overlap between stigma and anthers within an inflorescence. Large inflorescences attract more pollinators, potentially enhancing reproductive success by increasing pollen import and export (Schemske, 1980; Queller, 1983; Bell, 1985; Geber, 1985; Schmid-Hempel & Speiser, 1988; Klinkhamer & de Jong, 1990). However, large inflorescences also increase the opportunities for both geitonogamy and pollen discounting, so that the opportunity for between-flower interference increases with inflorescence size (Harder & Barrett, 1996). Consequently, the evolution of floral display size may represent a compromise between maximizing pollinator visitation and minimizing geitonogamy and pollen discounting (Klinkhamer & de Jong, 1993; Barrett et al., 1994; Holsinger, 1996; Snow et al., 1996).
Protandry may be particularly relevant to this compromise, because it often results in an inflorescence structure with female phase flowers positioned below male phase flowers (Bertin & Newman, 1993). Given the tendency of many insect pollinators to forage upwards through inflorescences (Galen & Plowright, 1988), protandry may enhance pollen export by reducing between-flower interference (Darwin, 1862; Harder et al., 2000). Furthermore, this enhanced pollen export should increase as floral display size increases, because between-flower interference should increase with floral display size. These effects of protandry on between-flower interference may decouple the benefits of large inflorescences from the consequences of geitonogamy and pollen discounting. Such a decoupling would provide a significant reproductive advantage through increased pollinator visitation and siring success.
Harder et al. (2000) demonstrated experimentally that dichogamy both reduced rates of self-fertilization and enhanced outcross siring success through reductions in geitonogamy and pollen discounting, respectively. Routley & Husband (2003) examined the influence of inflorescence size on this siring advantage and found a bimodal distribution with increased siring success with both small and large display sizes.
The length of stigmatic receptivity plays a key role in regulating the isolation of the male and female stages in dichogamous plants, and stigmatic receptivity can be influenced by both temperature and humidity.[14] Another study by Jersakova and Johnson, studied the effects of protandry on the pollination process of the moth pollinated orchid, ‘’Satyrium longicauda’’. They discovered that protandry tended to reduce the absolute levels of self-pollination and suggest that the evolution of protandry could be driven by the consequences of the pollination process for male mating success.[15] Another study that indicated that dichogamy might increase male pollination success was the study performed by Dai and Galloway.[16]
Zoology
Protandry
Protandry refers to organisms that are born male and at some point in their lifespan change sex to female. Protandrous animals include clownfish. Clownfish have a very structured society. In the Amphiprion percula species, there are zero to four individuals excluded from breeding and a breeding pair living in a sea anemone. Dominance is based on size, the female being the largest and the male being the second largest. The rest of the group is made up of progressively smaller non-breeders, which have no functioning gonads.[3] If the female dies, the male gains weight and becomes the female for that group. The largest non-breeding fish then sexually matures and becomes the male of the group.[4]Other examples of protandrous animals include:
- The ctenophore Coeloplana gonoctena. In this organism the females are bigger than the males and are only found during the summer. In contrast males are found year round.
- The flatworms Hymanella retenuova and Paravortex cardii.
- Laevapex fuscus, a gastropod, is described as being functionally protandric. The sperm matures in late winter and early spring, and the eggs mature in early summer, and copulation occurs only in June. This shows that males cannot reproduce until the females appear, thus why they are considered to be functionally protandric.[5]
[edit] Protogyny
Protogyny refers to organisms that are born female and at some point in their lifespan change sex to males. Common model organisms for this type of sequential hermaphroditism are wrasses. They are one of the largest families of coral reef fish and belong to the Labridae family. Wrasses are found around the world in all marine habitats and tend to bury themselves in sand at night or when they feel threatened.[6] In wrasses, the larger of the two fish is the male, while the smaller is the female. In most cases, females and immature have a uniform color while the male has the terminal bicolored phase.[7] Large males hold territories and try to pair spawn while small to mid-size initial-phase males live with females and group spawn.[8] In other words, both the initial and terminal phase males can breed; they differ however in the way they do it.In the California Sheephead (Pimelometopon pulchrum), a type of wrasse, when the female changes to male, the ovaries degenerate and spermatogenic crypts appear in the gonads. The general structure of the gonads remains ovarian after the transformation and the sperm is transported through a series of ducts on the periphery of the gonad and oviduct. Here sex change is age dependant. For example, the California sheephead stays a female for four years before changing sex.[7]
Other examples of protogynous organisms include:
- The isopods Cyathura polita and C. carinata
- The tanaidacean Heterotanais oerstedi.
- The echinoderms, Asterina pancerii and A. gibbosa are also protogynous and they brood their young.
- Protogyny sometimes occurs in the frog Rana temporaria, where old females sometimes change to males.[5]
[edit] Ultimate causes
Ghiselin proposed three models for hermaphroditism in 1969 in his paper titled “The evolution of hermaphroditism among animals”. The ‘’low-density model’’ states that individuals have characteristics that reduce the opportunity for mating; this model cannot be applied to sequential hermaphroditism. The ‘’gene dispersal model’’ is based on the idea that limitations on dispersal may influence population structure or genetical environment and it can be separated into two versions: the inbreeding version and the sampling-error version. This theory of gene dispersal can be applied to sequential hermaphrodites, especially the inbreeding version. The inbreeding version is based upon the fact that both protandry and protogyny help prevent inbreeding in plants and thus one can make the same assumption that in animals it works by reducing the probability of this occurring among siblings. The sampling-error version is based on the reality that the genetical environment is influenced by genetic drift and similar phenomena in small populations. The two aspects of these hypotheses influenced by hermaphroditism, that is inbreeding and sampling-error, result in the same thing, reduction of genetic variability. In other words being a hermaphrodite would increase genetic variability and thus be considered advantageous to the organism. This theory of gene dispersal can be applied to sequential hermaphrodites, especially the inbreeding version. Lastly, the ‘’size-advantage model’’ states that reproductive functions are carried out better if the individual is a certain size/age. Assuming that the reproductive functions of one sex are better performed at a certain size, then an organism would assume the sex that its size allows to perform the best. This would increase its reproductive potential and fitness. For example, eggs are larger than sperm, thus if you are a big you are able to make more eggs so being female when big is advantageous, however the size advantage relationship is really not as simple as the example just mentioned, but it allows for a better understanding of it.[9]In most ectotherms body size and female fecundity are positively correlated.[1] This supports Ghiselin’s size-advantage model, which is still widely accepted today. Kazancioglu and Alonzo (2010) performed the first comparative analysis of sex change in Labridae. Their analysis supports the size-advantage model by Ghiselin and suggest that sequential hermaphroditism is correlated to the size-advantage. They determined that dioecy was less likely to occur when the size advantage is stronger than other advantages[10]
Warner suggests that selection for protandry may occur in populations where female fecundity is augmented with age and individuals mate randomly. Selection for protogyny may occur where there are traits in the population that depress male fecundity at early ages (territoriality, mate selection or inexperience) and when female fecundity is decreased with age, the latter seems to be rare in the field.[1] An example of territoriality favoring protogyny occurs when there is a need to protect their habitat and being a large male is advantageous for this purpose. In the mating aspect, a large male has a higher chance of mating, while this has no effect on the female mating fitness.[10] Thus, he suggests that female fecundity has more impact on sequential hermaphroditism that the age structures of the population.[1]
The size-advantage model predicts that sex change would only be absent if the relationship between size/age with reproductive potential is identical in both sexes. With this prediction one would assume that hermaphroditism is very common, but this is not the case. Sequential hermaphroditism is very rare and according to scientists this is due to some cost that decreases fitness in sex changers as opposed to those who don’t change sex. Kazanciglu and Alonzo confirmed this in 2009. They found that the costs of changing sex only favored dioecy when the cost was very large but that some groups favored hermaphroditism. This indicates that the cost of sex change does not explain the rarity of sequential hermaphroditism by itself.[11]
[edit] Proximate causes
Many studies have focused on the proximate causes of sequential hermaphroditism. The role of aromatase has been widely studied in this area. Aromatase is an enzyme that controls the androgen/estrogen ratio in animals by catalyzing the conversion of testosterone into oestradiol, which is irreversible. It has been discovered that the aromatase pathway mediates sex change in both directions.[12] Many studies also involve understanding the effect of aromatase inhibitors on sex change. One such study was performed by Kobayashi et al. In their study they tested the role of estrogens in male three-spot wrasses (Halichoeres trimaculatus). They discovered that fish treated with aromatase inhibitors showed decreased gonodal weight, plasma estrogen level and spermatogonial proliferation in the testis as well as increased androgen levels. Their results suggest that estrogens are important in the regulation of spermatogenesis in this protogynous hermaphrodite.[13][edit] Botany
[edit] Flowering plants
In the context of the plant sexuality of flowering plants (angiosperms), there are two forms of dichogamy: protogyny—female function precedes male function—and protandry—male function precedes female function.Historically, dichogamy has been regarded as a mechanism for reducing inbreeding (e.g., Darwin, 1862). However, a survey of the angiosperms found that self-incompatible (SI) plants, which are incapable of inbreeding, were as likely to be dichogamous as were self-compatible (SC) plants (Bertin, 1993). This finding led to a reinterpretation of dichogamy as a more general mechanism for reducing the impact of pollen-pistil interference on pollen import and export (reviewed in Lloyd & Webb, 1986; Barrett, 2002). Unlike the inbreeding-avoidance hypothesis, which focused on female function, this interference-avoidance hypothesis considers both gender functions.
In many hermaphroditic species, the close physical proximity of anthers and stigma makes interference unavoidable, either within a flower or between flowers on an inflorescence. Within-flower interference, which occurs when either the pistil interrupts pollen removal or the anthers prevent pollen deposition, can result in autonomous or facilitated self-pollination (Lloyd & Webb, 1986; Lloyd & Schoen, 1992). Between-flower interference results from similar mechanisms, except that the interfering structures occur on different flowers within the same inflorescence and it requires pollinator activity. This results in geitonogamous pollination, the transfer of pollen between flowers of the same individual (Lloyd & Schoen, 1992; de Jong et al., 1993). In contrast to within-flower interference, geitonogamy necessarily involves the same processes as outcrossing: pollinator attraction, reward provisioning, and pollen removal. Therefore, between-flower interference not only carries the cost of self-fertilization (inbreeding depression; Charlesworth & Charlesworth, 1987; Husband & Schemske, 1996), but also reduces the amount of pollen available for export (so-called "pollen discounting"; Harder & Wilson, 1998]). Because pollen discounting diminishes outcross siring success, interference avoidance may be an important evolutionary force in floral biology (Harder & Barrett, 1995, 1996; Harder & Wilson, 1998; Barrett, 2002).
Dichogamy may reduce between-flower interference by minimizing the temporal overlap between stigma and anthers within an inflorescence. Large inflorescences attract more pollinators, potentially enhancing reproductive success by increasing pollen import and export (Schemske, 1980; Queller, 1983; Bell, 1985; Geber, 1985; Schmid-Hempel & Speiser, 1988; Klinkhamer & de Jong, 1990). However, large inflorescences also increase the opportunities for both geitonogamy and pollen discounting, so that the opportunity for between-flower interference increases with inflorescence size (Harder & Barrett, 1996). Consequently, the evolution of floral display size may represent a compromise between maximizing pollinator visitation and minimizing geitonogamy and pollen discounting (Klinkhamer & de Jong, 1993; Barrett et al., 1994; Holsinger, 1996; Snow et al., 1996).
Protandry may be particularly relevant to this compromise, because it often results in an inflorescence structure with female phase flowers positioned below male phase flowers (Bertin & Newman, 1993). Given the tendency of many insect pollinators to forage upwards through inflorescences (Galen & Plowright, 1988), protandry may enhance pollen export by reducing between-flower interference (Darwin, 1862; Harder et al., 2000). Furthermore, this enhanced pollen export should increase as floral display size increases, because between-flower interference should increase with floral display size. These effects of protandry on between-flower interference may decouple the benefits of large inflorescences from the consequences of geitonogamy and pollen discounting. Such a decoupling would provide a significant reproductive advantage through increased pollinator visitation and siring success.
Harder et al. (2000) demonstrated experimentally that dichogamy both reduced rates of self-fertilization and enhanced outcross siring success through reductions in geitonogamy and pollen discounting, respectively. Routley & Husband (2003) examined the influence of inflorescence size on this siring advantage and found a bimodal distribution with increased siring success with both small and large display sizes.
The length of stigmatic receptivity plays a key role in regulating the isolation of the male and female stages in dichogamous plants, and stigmatic receptivity can be influenced by both temperature and humidity.[14] Another study by Jersakova and Johnson, studied the effects of protandry on the pollination process of the moth pollinated orchid, ‘’Satyrium longicauda’’. They discovered that protandry tended to reduce the absolute levels of self-pollination and suggest that the evolution of protandry could be driven by the consequences of the pollination process for male mating success.[15] Another study that indicated that dichogamy might increase male pollination success was the study performed by Dai and Galloway.[16]
Huwebes, Oktubre 6, 2011
Accelerating universe/Accelerating expansion of the cosmos
Saul Perlmutter(right)
Brian P. Schmidt(middle)
Adam G. Riess(left)
These Nobel prize for physics winners contribute for the accelerating expansion of the universe by observing a supernovae
The accelerating universe is the observation that the universe appears to be expanding at an increasing rate, which in formal terms means that the cosmic scale factor a(t) has a positive second derivative,[1] implying that the velocity at which a given galaxy is receding from us should be continually increasing over time[2] (here the recession velocity is the same one that appears in Hubble's law; defining 'velocity' in cosmology is somewhat subtle, see Comoving distance#Uses of the proper distance for a discussion). In 1998, observations of Type Ia supernovae suggested that the expansion of the universe has been accelerating[3][4] since around redshift of z~0.5.[5] The 2006 Shaw Prize in Astronomy and the 2011 Nobel Prize in Physics were both awarded to Saul Perlmutter, Brian P. Schmidt, and Adam G. Riess for the 1998 discovery of the accelerating expansion of the Universe through observations of distant supernovae.[6][7]
In cosmology, the notion of an Accelerating Expansion of the cosmos is that the matter of the observed universe that is expanding outward from the big-bang is also expanding at an accelerating velocity.
Miyerkules, Oktubre 5, 2011
Blueshift
Redshift above, blueshift below
A blueshift is any decrease in wavelength (increase in frequency); the opposite effect is referred to as redshift. In visible light, this shifts the colour from the red end of the spectrum to the blue end. The term also applies when photons outside the visible spectrum (e.g. x-rays and radio waves) are shifted towards shorter wavelengths, as well as to shifts in the de Broglie wavelength of particles. Blueshift is most commonly caused by relative motion towards the observer, described by the Doppler effect. An observer in a gravity well will also see infalling radiation gravitationally blueshifted, described by General Relativity in the same way as gravitational redshift. In a contracting universe, cosmological blueshift would be observed; the expanding universe gives a cosmological redshift, and the expansion is observed to be accelerating.
Redshift
Redshift above, blueshift below
In physics (especially astrophysics), redshift happens when light seen coming from an object is proportionally increased in wavelength, or shifted to the red end of the spectrum. More generally, where an observer detects electromagnetic radiation outside the visible spectrum, "redder" amounts to a technical shorthand for "increase in electromagnetic wavelength" — which also implies lower frequency and photon energy in accord with, respectively, the wave and quantum theories of light.
Redshifts are attributable to the Doppler effect, familiar in the changes in the apparent pitches of sirens and frequency of the sound waves emitted by speeding vehicles; an observed redshift due to the Doppler effect occurs whenever a light source moves away from an observer. Cosmological redshift is seen due to the expansion of the universe, and sufficiently distant light sources (generally more than a few million light years away) show redshift corresponding to the rate of increase of their distance from Earth. Finally, gravitational redshifts are a relativistic effect observed in electromagnetic radiation moving out of gravitational fields. Conversely, a decrease in wavelength is called blue shift and is generally seen when a light-emitting object moves toward an observer or when electromagnetic radiation moves into a gravitational field.
Although observing redshifts and blue shifts have several terrestrial applications (e.g., Doppler radar and radar guns),[1] redshifts are most famously seen in the spectroscopic observations of astronomical objects.[2]
A special relativistic redshift formula (and its classical approximation) can be used to calculate the redshift of a nearby object when spacetime is flat. However, many cases such as black holes and Big Bang cosmology require that redshifts be calculated using general relativity.[3] Special relativistic, gravitational, and cosmological redshifts can be understood under the umbrella of frame transformation laws. There exist other physical processes that can lead to a shift in the frequency of electromagnetic radiation, including scattering and optical effects; however, the resulting changes are distinguishable from true redshift and not generally referred as such (see section on physical optics and radiative transfer).
Lunes, Oktubre 3, 2011
Fog bow
A fog bow is a similar phenomenon to a rainbow, however, as its name suggests, it appears as a bow in fog rather than rain. Because of the very small size of water droplets that cause fog—smaller than 0.05 millimetres (0.0020 in)—the fog bow has only very weak colors, with a red outer edge and bluish inner.
In many cases when the droplets are very small, fog bows appear white, and are therefore sometimes called white rainbows. This lack of color is a distinguishing feature from a glory, which has multiple pale colored rings caused by diffraction. When the droplets forming it are almost all of the same size the fog bow can have multiple inner rings, or supernumeraries, that are more strongly colored than the main bow. According to NASA:
In many cases when the droplets are very small, fog bows appear white, and are therefore sometimes called white rainbows. This lack of color is a distinguishing feature from a glory, which has multiple pale colored rings caused by diffraction. When the droplets forming it are almost all of the same size the fog bow can have multiple inner rings, or supernumeraries, that are more strongly colored than the main bow. According to NASA:
The fogbow's lack of colors is caused by the smaller water drops ... so small that the wavelength of light becomes important. Diffraction smears out colors that would be created by larger rainbow water drops ...[1]A fog bow seen in clouds, typically from an aircraft looking downwards, is called a cloud bow. Mariners sometimes call fog bows sea-dogs.
Mendel's laws
Mendel discovered that when crossing white flower and purple flower plants, the result is not a blend. Rather than being a mix of the two, the offspring was purple flowered. He then conceived the idea of heredity units, which he called "factors", one of which is a recessive characteristic and the other dominant. Mendel said that factors, later called genes, normally occur in pairs in ordinary body cells, yet segregate during the formation of sex cells. Each member of the pair becomes part of the separate sex cell. The dominant gene, such as the purple flower in Mendel's plants, will hide the recessive gene, the white flower. After Mendel self-fertilized the F1 generation and obtained the 3:1 ratio, he correctly theorized that genes can be paired in three different ways for each trait: AA, aa, and Aa. The capital "A" represents the dominant factor and lowercase "a" represents the recessive. (The last combination listed above, Aa, will occur roughly twice as often as each of the other two, as it can be made in two different ways, Aa or aA.)
Mendel stated that each individual has two factors for each trait, one from each parent. The two factors may or may not contain the same information. If the two factors are identical, the individual is called homozygous for the trait. If the two factors have different information, the individual is called heterozygous. The alternative forms of a factor are called alleles. The genotype of an individual is made up of the many alleles it possesses. An individual's physical appearance, or phenotype, is determined by its alleles as well as by its environment. An individual possesses two alleles for each trait; one allele is given by the female parent and the other by the male parent. They are passed on when an individual matures and produces gametes: egg and sperm. When gametes form, the paired alleles separate randomly so that each gamete receives a copy of one of the two alleles. The presence of an allele doesn't promise that the trait will be expressed in the individual that possesses it. In heterozygous individuals the only allele that is expressed is the dominant. The recessive allele is present but its expression is hidden.
Mendel summarized his findings in two laws; the Law of Segregation and the Law of Independent Assortment.
More precisely the law states that when any individual produces gametes, the copies of a gene separate so that each gamete receives only one copy (allele). A gamete will receive one allele or the other. The direct proof of this was later found following the observation of meiosis by two independent scientists, the German botanist, Oscar Hertwig in 1876, and the Belgian zoologist, Edouard Van Beneden in 1883. In meiosis, the paternal and maternal chromosomes get separated and the alleles with the traits of a character are segregated into two different gametes.
Independent assortment occurs during meiosis I in eukaryotic organisms, specifically metaphase I of meiosis, to produce a gamete with a mixture of the organism's maternal and paternal chromosomes. Along with chromosomal crossover, this process aids in increasing genetic diversity by producing novel genetic combinations.
Of the 46 chromosomes in a normal diploid human cell, half are maternally-derived (from the mother's egg) and half are paternally-derived (from the father's sperm). This occurs as sexual reproduction involves the fusion of two haploid gametes (the egg and sperm) to produce a new organism having the full complement of chromosomes. During gametogenesis—the production of new gametes by an adult—the normal complement of 46 chromosomes needs to be halved to 23 to ensure that the resulting haploid gamete can join with another gamete to produce a diploid organism. An error in the number of chromosomes, such as those caused by a diploid gamete joining with a haploid gamete, is termed aneuploidy.
In independent assortment the chromosomes that end up in a newly-formed gamete are randomly sorted from all possible combinations of maternal and paternal chromosomes. Because gametes end up with a random mix instead of a pre-defined "set" from either parent, gametes are therefore considered assorted independently. As such, the gamete can end up with any combination of paternal or maternal chromosomes. Any of the possible combinations of gametes formed from maternal and paternal chromosomes will occur with equal frequency. For human gametes, with 23 pairs of chromosomes, the number of possibilities is 223 or 8,388,608 possible combinations.[3] The gametes will normally end up with 23 chromosomes, but the origin of any particular one will be randomly selected from paternal or maternal chromosomes. This contributes to the genetic variability of progeny.
Mendel stated that each individual has two factors for each trait, one from each parent. The two factors may or may not contain the same information. If the two factors are identical, the individual is called homozygous for the trait. If the two factors have different information, the individual is called heterozygous. The alternative forms of a factor are called alleles. The genotype of an individual is made up of the many alleles it possesses. An individual's physical appearance, or phenotype, is determined by its alleles as well as by its environment. An individual possesses two alleles for each trait; one allele is given by the female parent and the other by the male parent. They are passed on when an individual matures and produces gametes: egg and sperm. When gametes form, the paired alleles separate randomly so that each gamete receives a copy of one of the two alleles. The presence of an allele doesn't promise that the trait will be expressed in the individual that possesses it. In heterozygous individuals the only allele that is expressed is the dominant. The recessive allele is present but its expression is hidden.
Mendel summarized his findings in two laws; the Law of Segregation and the Law of Independent Assortment.
[edit] Law of Segregation (The "First Law")
The Law of Segregation states that every individual possesses a pair of alleles for any particular trait and that each parent passes a randomly selected copy (allele) of only one of these to its offspring. The offspring then receives its own pair of alleles for that trait. Whichever of the two alleles in the offspring is dominant determines how the offspring expresses that trait (e.g. the color of a plant, the color of an animal's fur, the color of a person's eyes).More precisely the law states that when any individual produces gametes, the copies of a gene separate so that each gamete receives only one copy (allele). A gamete will receive one allele or the other. The direct proof of this was later found following the observation of meiosis by two independent scientists, the German botanist, Oscar Hertwig in 1876, and the Belgian zoologist, Edouard Van Beneden in 1883. In meiosis, the paternal and maternal chromosomes get separated and the alleles with the traits of a character are segregated into two different gametes.
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[edit] Law of Independent Assortment (The "Second Law")
The Law of Independent Assortment, also known as "Inheritance Law" states that separate genes for separate traits are passed independently of one another from parents to offspring. That is, the biological selection of a particular gene in the gene pair for one trait to be passed to the offspring has nothing to do with the selection of the gene for any other trait. More precisely the law states that alleles of different genes assort independently of one another during gamete formation. While Mendel's experiments with mixing one trait always resulted in a 3:1 ratio (Fig. 1) between dominant and recessive phenotypes, his experiments with mixing two traits (dihybrid cross) showed 9:3:3:1 ratios (Fig. 2). But the 9:3:3:1 table shows that each of the two genes are independently inherited with a 3:1 phenotypic ratio. Mendel concluded that different traits are inherited independently of each other, so that there is no relation, for example, between a cat's color and tail length. This is actually only true for genes that are not linked to each other.Independent assortment occurs during meiosis I in eukaryotic organisms, specifically metaphase I of meiosis, to produce a gamete with a mixture of the organism's maternal and paternal chromosomes. Along with chromosomal crossover, this process aids in increasing genetic diversity by producing novel genetic combinations.
Of the 46 chromosomes in a normal diploid human cell, half are maternally-derived (from the mother's egg) and half are paternally-derived (from the father's sperm). This occurs as sexual reproduction involves the fusion of two haploid gametes (the egg and sperm) to produce a new organism having the full complement of chromosomes. During gametogenesis—the production of new gametes by an adult—the normal complement of 46 chromosomes needs to be halved to 23 to ensure that the resulting haploid gamete can join with another gamete to produce a diploid organism. An error in the number of chromosomes, such as those caused by a diploid gamete joining with a haploid gamete, is termed aneuploidy.
In independent assortment the chromosomes that end up in a newly-formed gamete are randomly sorted from all possible combinations of maternal and paternal chromosomes. Because gametes end up with a random mix instead of a pre-defined "set" from either parent, gametes are therefore considered assorted independently. As such, the gamete can end up with any combination of paternal or maternal chromosomes. Any of the possible combinations of gametes formed from maternal and paternal chromosomes will occur with equal frequency. For human gametes, with 23 pairs of chromosomes, the number of possibilities is 223 or 8,388,608 possible combinations.[3] The gametes will normally end up with 23 chromosomes, but the origin of any particular one will be randomly selected from paternal or maternal chromosomes. This contributes to the genetic variability of progeny.
Mendelian inheritance
Mendelian inheritance (or Mendelian genetics or Mendelism) is a scientific description of how hereditary characteristics are passed from parent organisms to their offspring; it underlies much of genetics. This theoretical framework was initially derived from the work of Gregor Johann Mendel published in 1865 and 1866 which was re-discovered in 1900; it was initially very controversial. When Mendel's theories were integrated with the chromosome theory of inheritance by Thomas Hunt Morgan in 1915, they became the core of classical genetics.
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